Date of Award

5-2025

Document Type

Thesis

Degree Name

Master of Science (MS)

Department

Mathematical Sciences

Committee Member

Keisha Cook (Co-Advisor)

Committee Member

Matthew Macauley (Co-Advisor)

Committee Member

Jennifer Mason

Abstract

DNA can be damaged through both internal and external sources. Therefore, cells have created methods to repair DNA damage. In Escherichia coli, the system responsible for DNA repair is termed the SOS response. This system consists of more than 50 genes and contains three main repair pathways: nucleotide excision repair, translesion synthesis, and homologous recombination. The response is initiated when DNA lesions result in the accumulation of single-stranded DNA (ssDNA). The protein RecA is activated by binding to ssDNA and is then denoted RecA*. RecA* assists in the auto-cleavage of LexA which is the primary repressor protein involved in the SOS system. Inactivation of LexA allows for the transcription of operons, such as umuDC, uvrAB, recX, recN, recX, lexA, dinI, and yebG. In this study, we model the behavior of these genes and their gene products during the SOS response using a stochastic model and a Boolean model. In the stochastic model, we observe oscillations in some of the components of the system as well as two distinct responses: one in which DNA is repaired and the other where DNA is not repaired within 330 minutes. We find that the Boolean model exhibits two fixed points when the SOS system is induced which also suggests an oscillatory behavior and/or bistability. We hypothesize that these observations are due to 1) stalling/restarting the replication fork and 2) proteins inhibiting RecA*. In comparing our model to previous experimental results, we observe that it matches these data.

Author ORCID Identifier

https://orcid.org/0000-0002-1618-9587

Share

COinS
 
 

To view the content in your browser, please download Adobe Reader or, alternately,
you may Download the file to your hard drive.

NOTE: The latest versions of Adobe Reader do not support viewing PDF files within Firefox on Mac OS and if you are using a modern (Intel) Mac, there is no official plugin for viewing PDF files within the browser window.