Date of Award

8-2025

Document Type

Thesis

Degree Name

Master of Science (MS)

Department

Animal and Veterinary Sciences

Committee Chair/Advisor

Dr. Susan Duckett

Committee Member

Dr. William Bridges

Committee Member

Dr. Elliot Jesch

Abstract

Background

Satellite cells (SCs) are myogenic precursor cells that play a crucial role in muscle growth, and their activity can be influenced by nutrient supplementation. Muscle growth can also be stimulated using growth-promoting hormones; however, there are no approved growth hormones for use in sheep in the United States (Qaid & Abdoun, 2022). This limitation highlights a need to identify alternative compounds that would serve as nutrient supplements to enhance muscle growth and thereby improve meat production. Amino acid derivatives such as guanidinoacetic acid (GAA) and 5-hydroxytryptophan (5-HTP), along with the polyphenol quercetin, are naturally derived compounds that have shown to positively influence muscle growth (Chandran et al., 2012; Michiels et al., 2012; Wang et al., 2022). It is therefore important to investigate how these compounds may contribute to skeletal muscle growth in sheep.

Methods

Satellite cells were cultured in a Corning 24-well plate (45,000/well). The media was supplemented with different levels (0, 12.5, 25, and 50 μmol/L) of 5-HTP, GAA, or quercetin. Cells were counted after day 4 for proliferation and differentiation. For the in vivo study, 15 lambs were used (33 kg + 6 kg BW) with five lambs per group assigned to one of the treatments: control, GAA, or 5-HTP. Lambs were individually fed for 28 days, and supplements were administered at a dose of 2.5 mg/kg body weight via the sublingual route before feeding. At the end of the study, a muscle biopsy was carried out from the longissimus muscle at the 12th/13th rib, and real-time ultrasound measurements of ribeye area (REA) were recorded. The muscle samples were subsequently processed for mRNA sequencing to assess gene expression profiles.

Results

5-HTP (25 μmol/L) increased satellite cell number (P< 0.05), and 5-HTP (12.5 and 25 μmol/L) increased proliferation of SC (P< 0.05). There was no effect of 5-HTP on SC differentiation (P>0.05). Quercetin (12.5 and 25 μmol/L) also increased SC proliferation when compared with 0 μmol/L (P< 0.05), with no effect on SC differentiation (P>0.05). GAA altered SC differentiation in a dose-dependent manner, with 12.5 and 25 μmol/L differing from 0 μmol/L (P < 0.05), but showed no effect on SC proliferation (P>0.05). For lamb characteristics, GAA or 5-HTP did not influence (P>0.05) average daily gain, dry matter intake, or ribeye area compared to the control. miRNA qPCR showed that GAA nor 5-HTP did not alter miR-133a-3p and miR-133a-5p (P>0.05) expression compared with the control. mRNA sequencing showed that GAA supplementation altered gene expression (Padj < 0.05) compared to the control or 5-HTP. However, 5-HTP did not affect (P>0.05) gene expression compared to the control.

Conclusion

These data suggest that nutrient supplementation could influence satellite proliferation and differentiation in vitro, which could enhance muscle mass accretion. Short-term supplementation of GAA or 5-HTP to finishing lambs did not alter growth performance, carcass REA or miRNA expression; however, GAA supplementation did alter gene expression of four genes that are involved in muscle growth.

Available for download on Monday, August 31, 2026

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